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The Forbidden Tower (pub. 1977) is the story of the creation of the "heretical" circle of telepaths who, choosing to work outside of the Towers of Darkover, challenge the two most strongly held beliefs that underlie the Tower system - that a Keeper must be essentially asexual, and that only the members of the ruling Comyn caste have enough laran to work in the Towers.

The novel begins where The Spell Sword left off - with the clearing of the catmen and the planned union of Damon Ridenow to Ellemir Alton, and of Terran Andrew Carr to Callista Alton, formerly a Keeper of Arilinn under Leonie Hastur. Much of the main plot of the novel deals with the fusion of these two couples into a fourway bond, linked telepathically, emotionally and sexually. There are two main obstacles to this, and MZB deals with both in great detail. First, the realisation that not only has Callista been conditioned to have no sexual awareness or response, but that early in the training, Leonie performed a kind of psychic neutering on her, so that it would be impossible for her conditioning to ever be undone. Second, the painful misunderstandings and problems of culture shock brought about by the differences between Terran sexual mores and those found in a society of telepaths. In order to overcome the first, Damon must engage in the dangerous discipline of timesearch to find clues to a centuries-old tradition that could restore Callista's frozen sexuality. And only endless love and patience can overcome the second.

At the end of the novel, the four of them, fully bonded, are faced with a telepathic duel to prove Damon's right to namr himself Keeper and to direct the way his Tower will operate according to his own conscience and not the laws of Arilinn.

While largely focused on deconstructing the rigid role of Keeper and the assumption that only the Comyn can be effective telepaths, many of the Darkovan attitudes toward sexuality are clarified through the exploration of the differences between Terran and Darkovan sexual culture.

Darkovan society is to some degree polyamorous, and despite the strongly patriarchal nature of family relationship, women appear to have some sexual autonomy, but on strict class lines. As well, women must be discreet, and if unmarried, must be careful about pregnancy. The greatest shame seems to lie in bearing a child who has no acknowledged father. Some of the contradictions are shown in this account Ellemir gives to Callista about her sexual experience:
“It was that winter,” said Ellemir. “Dorian begged me to come and spend the winter with her; she was lonely, and already pregnant, and had made few friends of the mountain women. Father gave me leave to go. And later in the spring, when Dorian grew heavy, so it was no pleasure to her to share his bed, Mikhail and I had grown to be such friends that I took her place there.” She giggled a little, reminiscently. Callista said, startled, “You were no more than fifteen!” Ellemir answered, laughing, “That is old enough to marry; Dorian had been no more. I would have been married, had Father not wanted me to stay home and keep his house!” Again Callista felt the cruel envy, the sense of desperate alienation. How simple it had been for Ellemir, and how right! And how different for her! “Were there others?” Ellemir smiled in the darkness. “Not many. I learned there that I liked lying with men, but I did not want to be gossiped about as they whisper scandal about Sybil-Mhari—you have heard that she takes lovers from Guardsmen or even grooms—and I did not want to bear a child I would not be allowed to rear, though Dorian pledged that if I gave Mikhail a child she would foster it. And I did not want to be married off in a hurry to someone I did not like, which I knew Father would do if there was scandal."
There is some indication, however, that the circumstances in which women may engage in pre- or extra-marital sex are partly for the convenience of men. There is a reluctance among Darkovan women to engage in sex during pregnancy. As Callista explains, “Biologically, no pregnant animal desires sex; most will not endure it. If your women have been culturally conditioned to accept it as the price of retaining a husband's sexual interest, I can only say I am sorry for them! Would you demand it of me after I had ceased to take pleasure in it?”

While a man may take a concubine or mistress at his pleasure, and it is expected that he will do so if his wife is unavailable or unable to provide sex, it is considered not quite proper if he brings into the household a woman who is not acceptable to his wife. The kinswoman of one's wife is traditionally one of the more acceptable choices in such circumstances.
"This is our custom. If you were one of us, it would be taken for granted that my sister and I should… should share in this way. Even if things were — as they should be between us, if there was a time when I was ill, or pregnant, or simply not… not wanting you… It is very old, this custom. You have heard me sing the Ballad of Hastur and Cassilda? Even there, even in the ballad, it speaks of how Camilla took the place of her breda in the arms of the God, and so died when he was set upon. It was so that the Blessed Cassilda survived the treachery of Alar, to bear the child of the God…”
There is also a sense that men's desire should not be allowed to go unfulfilled, and that women are responsible for seeing to this when they arouse a man's sexual interest.
In both The Spell Sword and The Forbidden Tower, Callista tells Andrew that she is responsible for the fulfilment of desire he feels toward her. “I have been taught that it is… shameful to arouse a desire I will not satisfy."

Despite the suggestion of some choice in sexual expression, it is also clear that women, like children must always be under wardship - except, of course, for the Keepers. Women in the Towers are under the wardship of their Keepers - at one point, Leonie states that it is her responsibility to find suitable marriages for women who have given oath to her as Keeper and have worked in the Tower (this does not include young women who are sent to the Towers for a few years training in the use of laran) if they later choose to leave. Women outside the Towers are seen always as under the authority of father, husband, brother, or other kinsman.

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